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Electrostatic interactions (including hydration) are vital to the structure and function of proteins (Hayes and Kollman, 1976, Honig and Nicholls, 1995, Sham et al., 1998, Warshel and Aqvist, 1991, Warshel, 1991).A particularly straightforward method for probing internal electrostatics is the use of a chromophore whose wavelength is sensitive to electric fields. 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