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goat-antelope, or caprid, group is known from as early as the Miocene, when members of the group resembled the modern serow in their general body form. The group did not reach its greatest diversity until the recent ice ages, when many of its members became specialised for marginal, often extreme, environments: mountains, deserts, and the subarctic region.
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Territorial behaviour occurs in many animals and is especially widespread in mammals. Both visual and chemical signals may be used to advertise the territory to other animals.
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